The box can only be checked and unchecked upon creation of the Item Receipt. Once the Item Receipt is saved, there will be no other way to have it checked or unchecked.
Solution:
1. Edit it and manually assign the Landed Cost on the line Item level, instead.

Note: Once it is save, the GL Impact of the Item Receipt will now reflect the Landed Cost.
OR
2. Recreate the Item Receipt.
The Landed Cost fields are grayed out because the LANDED COST PER LINE box under Items & Expenses tab is marked.
The Landed Cost Per Line box can no longer be unmarked because the Item Receipt has already been saved.